The CIA's London station captures former CTU agent Jack Bauer. Jack remains silent under Station Chief Steve Navarro's questioning, only wavering when Agent Kate Morgan asks if he's looking for someone. Kate asks Navarro if she can take over the case, partially to atone for a recent disgrace (her husband turned out to be selling secrets to the Chinese), but Jack is turned over to a separate branch called Special Activities. This turns out to have been his plan all along, as said branch is holding Chloe O'Brian captive. With help from his friend Belcheck, he escapes Kate's pursuit and exfiltrates her. President James Heller, his daughter Audrey and her husband, White House Chief of Staff Mark Boudreau, are in London, amidst strident public opposition, to negotiate for renewal of the U.S. military lease of Diego Garcia, a British overseas territory in the Indian Ocean on which the U.S. unmanned aerial vehicles are stationed. Mark has been keeping track of progress on Jack's capture and is concerned he might come for the President and Audrey. Meanwhile, remotely piloting an armed drone from RAF Lower Heyford, Lt. Chris Tanner provides aerial reconnaissance to a U.S. military convoy in Afghanistan. The convoy stops and Tanner scans the area with the drone, but soon loses control of the drone. The drone fires at the convoy, and, despite Tanner's warnings, four soldiers are killed, including two British officers.
Chloe is a member of a hacker group, Open Cell, dedicated to exposing government secrets and led by Adrian Cross. Jack has business with them as he wants to track one of their members, Derrick Yates, a hacker now gone rogue. Derrick is shown to have been involved in hacking the drone that fired on the convoy. Tanner is questioned by his superiors, as his personal computer logs show that every command sent to the drone originated from his console. Kate demands to be re-instated as a field agent and to take over the hunt for Bauer, which Navarro allows, despite opposition from field agent Erik Ritter. President Heller shows signs of dementia, which worries Audrey and Mark, not only for his future as President but because he wishes to address the Parliament of the United Kingdom directly. With the help of Open Cell, Chloe and Jack track Derrick to drug dealer Basher's apartment. They are interrupted by the CIA; during the resulting stand-off, Derrick and his girlfriend escape with the device. Derrick expresses skepticism about his buyer continuing with the project, so his girlfriend kills him, steals the device, and calls her the buyer, and the mastermind of the drone attack.
After finding Yates's body, Jack realizes his girlfriend has the device. She escapes Jack in a pursuit on the London Underground, but Chloe identifies her as Simone Al-Harazi, daughter of international terrorist Margot Al-Harazi. Jack berates Chloe for missing Simone, but Chloe confesses that she saw what she thought was her husband Morris and son Prescott, who were killed in an attack she believes was intended for her. They return to Open Cell, where they hear about Tanner. Jack recruits Cross to help him get into the U.S. Embassy using a fake ID so that he can question Tanner. Kate and Ritter have taken Basher and his people hostage in the aftermath of the shooting. Navarro orders them to stand down, but she kidnaps Basher and forces him to divulge that Tanner is involved in the plot. Afterwards, Erik apologizes to Kate for his earlier dismissal of her abilities. After Simone returns to Margot's base of operations, Margot decides to use drones to attack ten British targets. However, the plan is delayed when special equipment is needed to activate the device. Jack attempts to enter the embassy, but Cross purposely botches the ID to get him arrested.
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