Summary: Prologue: In the Temple, Benjamin Sisko senses a change coming, though the Prophets won't tell him or don't know what it is. He is ready.
As they enter the wormhole and return to the Alpha Quadrant, Vaughn reflects on their mission, but is so deeply depressed about Ruriko and Prynn that he can't even appreciate what they accomplished. They exit the anomaly to find numerous Cardassian warships waiting for them, weapons armed, jamming contact with DS9. Gul Macet tells them to stand down and prepare to be boarded; Vaughn slumps in defeat and agrees. He tells Nog to cloak the ship, and they dart away.
Prynn does some hairy maneuvering, but at the last second, the Cardassians catch them with their sensor sweep; regardless, they get a message from Kira telling them to stand down. On screen, Kira and Akaar ask Vaughn and Bashir personal questions to confirm their identity; once they prove it's really them, they welcome them home--then say that the Cardassians will be examining the rest of the crew.
After the Cardassians inspect them, they're greeted by Kira, who is ecstatic/bewildered that Jake--and Opaka--are with the Defiant crew.
Kira debriefs the Defiant's senior crew with General Cyl from the Trill Defense Ministry. Dax appears uneasy to see Cyl there; one of Dax's previous hosts had been the mother of a woman who became a host of the Cyl symbiont. Kira explains that Shakaar was assassinated by Gard, who was infested by a parasite. One of the parasites had killed (Ezri's former host) Audrid Dax's husband, and the Trill Symbiosis Commission had covered it up--the lies of which lead to an estrangement between Audrid and her daughter Neema--Cyl's prior host. Kira says the parasites share a group consciousness, but can also act independently. Their numbers seem to be low, or they're slow to spread. Their goal appears to be the destruction of Trill. They've found 6 other infected people onboard, who are being held in medical stasis.
Akaar says a parasite attacked Gul Macet, but Cardassians appear to be immune--which may explain why Shakaar was trying to tank peace talks with them. He says a number of ships fled the station before it could be locked down, but of the ones that have been caught, no new infections were reported so far. However, there are still 107 Bajoran civilians that they can't account for.
Dr. Girani explains that the infected normally show a tiny barb on the back of their neck, but that in some cases, they have plastimasked the skin to hide it. They operate in hives controlled by a mother parasite that breeds inside her carrier, then transmits commands to her spawn telepathically. The parasite cannot likely be removed from the host after 3-4 weeks; though when the mother is killed, the solider parasites may be easily removed from their hosts. They can access a host's short-term memory, but not long-term. They can resist physical trauma, including low-level phaser fire.
Kira tells Vaughn to lead the Bajoran surface check with General Lenaris; Bashir will take over medical research with doctors Girani and Tarses; Shar and Nog are tasked with building scanning equipment; she tells Ezri to interview Gard.
After the meeting, Ezri tells Bashir that a comet passed near Trill ~100 years ago showing biosigns similar to those of the Trill. Audrid Dax and her husband Jayvin were on the landing party with 4 Federation men. They found a parasite in a frozen ice pool, and it infected Jayvin. He killed 3 of the Federation men, leaving only Christopher Pike alive. It chased after her for hours, screaming that the parasites were coming to obliterate the Trill. Eventually Pike subdued it, and they took it back to Trill. They found that the parasite had joined with the Vod symbiont, and decided to let it die.
The Trill covered up the incident (this was before any non-Trill knew about the symbionts) and sent ships to destroy the comet; only Pike and Audrid knew the truth. Later, her daughter Neema hacked into Audrid's logs and found out that Vod had died at her request, leading to their estrangement. Years later, Audrid wrote her a letter explaining everything, but Audrid died before they could fully reconcile. She says she knows Gard because he killed her.
Shar and his Zhavey return to his quarters. She says they all knew of his tezha with Thriss--intimacy outside of the shelthreth. Anichent and Dizhei embrace him, seeming not to hold him accountable for Thriss's death, but he can sense their resentment when he says he can't leave for Andor immediately. He tells them of the Yrythny eggs and their genetic key that could help the Andorians. Councillor zh'Thane says she's spoken with the Eveste Elders, and they've found 3 replacement zhen candidates...and 2 chans to replace Shar.
Ro stops in to ask Quark what rumors he's heard. He asks if she wants to run away with him, and she says she's still interested, but that she's not sure anymore, too busy to think about it. This makes him feel surprisingly better. Nog comes to greet him, and Quark is shocked that he didn't know the Defiant is back, what with all his sensors and contacts.
Kira visits Opaka in her quarters to get her up to speed. She tells her about the Ohalu book and how she was Attainted for it, then breaks down into tears. Opaka holds her until she falls asleep. She's unsure about what has happened to Bajor in her absence, not understanding how having it's own history exposed could result in such strict punishment. She sees Kira's strength as the future of Bajor.
Cyl tells Ezri that Neema respected Audrid, and never stopped researching the parasites. He says the government knows about them, but has their heads buried in the sand, hoping the Federation will make it go away. He's part of an unsanctioned shadow agency that keeps a watch out for the parasites--and so is Gard, though he went into business for himself with the assassination.
When Ezri meets Gard, she recognizes him as the man who stopped Joran Dax when his bad joining turned him into a serial killer. He fills her in on the rest of the story, saying that Shakaar sent out 7 coded messages to Bajor before he was killed. He also says the reason Trill hasn't done anything about the parasites is because there's an ingrained, primal fear of them. Ezri wants to tell Kira about the watch, and they agree, thinking maybe they'll finally get them under control.
Yevir approaches the station, having heard that Jake and Opaka have returned, and asks Kira to let him speak with them, insisting that they return to Bajor with him to bolster the people after the dire events. Kira agrees to let him aboard.
Shar decides that Anichent and Dizhei should return to Andor to search for 2 new partners; they resist only a little before agreeing. The councilor already pre-arranged transport for them, so they're off in short order. Prynn stops by to ask Shar to have lunch with her, upset at herself for not seeing her dad off to Bajor, but Shar can't even muster a polite "no."
Jake and Opaka decide to take Yevir up on his trip to Bajor, and they leave the next day. Wex decides to stay behind, though she tells Kira that Opaka asked her to. Opaka tells Jake that it was Wex's decision, but not to worry about it.
Ro spots Wex actually having a full on conversation with Taran'atar. Sure eavesdrops for a bit and learns that Taran'atar feels lost, without purpose, but can't return home because he wasn't given the option. When he calls out Ro for listening in, she apologizes and says she can hook them up at Quark's, and Wex accepts.
Vaughn already has a fully functioning base camp setup on Bajor. While Bowers gives him a report, he can't stop thinking about killing Ruriko, and is worried he might lose it.
After running some tests on the parasites, Bashir concludes that they didn't naturally evolve from symbionts: they were genetically engineered. Kira tells Ezri to prep the Defiant to head to Trill to present their findings to the TSC.
Gard tells Ro there were 5 places they believe Shakaar could've been infected, and he's willing to share them with her.
Shar apologies for being so awkward with Prynn earlier; she realizes that she had gone to talk to him because she's interested in him, and asks him to lunch again, to which he agrees.
Ro tells Kira that after speaking with Gard, she's narrowed down where Shakaar was infected to Minos Korva.
Vaughn feels himself in a downward spiral. He calls Prynn to say he's sorry, but she hangs up on him.
Quark bugs the connection between DS9 and the Trager and finds out about the parasites. He complains to Ro, but she says she couldn't tell him, and that she has Taran'atar watching his bar for extra protection. Then he gets a call from Rom who says Leeta is pregnant. Quark rudely congratulations them, then hangs up.
Security officers Kavi and Bennings search one of the unpowered docking rings. Kavi notices Bennings' light hasn't moved in a while, and when she turns around, he grins and grabs her.
General Lenaris tells Vaughn he's being sent to Ashalla monastery for R&R, per the doctor's orders. He reluctantly goes, knowing he truly is in a bad place, even if they did just turn up 11 new cases.
Shar and Prynn have lunch. She asks if Andorians ever date outside their species, and when he eventually realizes why, he's quite taken aback. She tells him no pressure, and he says he wouldn't be opposed to having lunch with her again some time, but he doesn't know what his personal life will entail yet. His zhavey sees Prynn touch his arm, and she turns her back on him.
Opaka is meditating when Vaughn arrives. She can sense that his pagh is greatly troubled, but decides not to press him until he's ready to unburden himself.
The O'Briens and Siskos' shuttle arrives at DS9, and after waiting in a security queue for a while, Kira welcomes them, and tells them that Jake is back. Miles starts to get nostalgic for working on the station, but promises Keiko it's only nostalgia.
Nog is dining at Quark's when one of the parasites runs through, setting off his vole traps. It jumps straight into Taran'atar's mouth, but he just chews it up and spits it out. Nog covers and says it was a pet of his, but only some of the patrons believe it.
Vaughn seeks out Opaka and finally tells her about Ruriko, Prynn, the Orb, etc. He finds he's still not sure what to do, and she tells him that the Tears of the Prophets are here.
The O'Briens and Siskos beam down to Kas's place and have a joyous reunion with Jake and Kas.
Ro heads to a meeting with Kira, where she overhears the major recommending a commendation for Ro's exemplary job to Akaar--who says now is not the time and tells her to come in out of the hallway. Ro turns around to see a Bajoran woman unhinge her jaw and vomit out a parasite, which latches onto Ro's face.
Shar is confiding to Nog that he can't stop thinking about Prynn when they hear Ro scream. Shar, in a bloodlust, attacks the queen, then rips the parasite out of Ro's mouth, smashing it into the ground. Nog calls for security, and they capture the queen with a stasis beam.
Kira contacts O'Brien and asks him to help Nog with the scanners; he agrees, and Keiko isn't happy. She tells him that the Interstellar Agricultural Aid Commission offered her a lead position to help restore Cardassia, and is angry that his job always comes before hers. He says they'll talk about it more when he gets back, but is not enthusiastic about raising their kids there. She decides she wants to takes the job.
Opaka has the Orb of Unity brought to Vaughn. The vision he sees is of himself being committed to a mental institution for shooting his wife with a revolver. The nurse introduces him to Benny Russell, who introduces him to the others, seemingly human versions of the DS9 crew. He enjoys talking to Benny, but doesn't feel like he can unburden himself to him.
Bashir presents his findings and says that the queen can't be implanted into another person because it's physiology changes after infecting someone, but the Ezri suggests maybe it would work in a Trill; Bashir says it's possible, but too risky. Ezri insists on trying it, then Cyl says he'll do it, before Gard eventually wins out.
Benny asks Eli if his has any kids, and says he has a baby on the way himself. He says he thought he was needed here but that time is slipping away and his family needs him. He tells Eli that he should keep trying to reconnect with his daughter, even if she resists.
Bashir surgically transports the queen into Gard. Shortly afterwards, the queen starts taking through him, telling Kira that it's already too late, that they will destroy Trill and take over the wormhole. Before she can take over Gard completely, Bashir removes the queen; Gard is OK, but the queen doesn't survive. Gard warns Kira that there's only one, that she sees Bajor, the Tears. They conclude that there's a single matriarch on Bajor controlling all the others. The parasites on the station start evacuating their hosts, and they also get word that skirmishes are breaking out on Bajor. Wex tries to tell Kira she can help, but she says she's too busy.
The Bajorans camped outside Kas's house show up at her house, armed with phasers, and force their way in.
The Orb Bearers break into the room with Elias and Opaka; they shoot Elias and take the Orb. Opaka tends to him, but his body, though still alive, is a soulless husk.
Elias has a nightmare about killing Ruriko; Benny wakes him up and tells him to stop blaming himself, seemingly able to read Eli's mind. He says he thought they had more time here, but says they have to leave because of what's happening outside. Benny says this is only one of the places he is, that he's here teaching and learning. He says as long as Eli thinks he killed his wife, he will stay in this place; he has to let his pain go and reconcile with Prynn. Eli sees that the security gate is open, and when Benny asks him to leave with him, he says he's ready.
Akaar orders Kira to confine herself to ops until he arrives at the station. She tells Ro to have Nog, O'Brien, and Dax to get the Defiant ready, and puts Ro in charge of the station. She is sure the matriarch is at the Ashalla Monastery.
Kassidy, Jake, and family have been kidnapped and taken to the monastery with Opaka and Vaughn, held under armed guard. Sulan feels Kassidy's stomach, and realizes she's having contractions.
O'Brien finds a convoluted way to beam Kira down to the monastery through the Defiant's cloak, where she's immediately attacked by infected Bajorans; however, Wex saves her, quickly and easily dispatching her assailants. Wex then liquefies; it's Odo. He had heard stories of a great healer that knew of the Ascendants, and decided to track her down to learn more. When he saw it was Opaka and Jake was with her, he stuck with them to make sure they got home safe. He kept his distance die to the crisis when they got back, and had hidden himself in Kira's belt holster when she beamed down. She tells him to do rescue the hostages and they split up before they can talk much.
Kira enters the once secret chamber where the Tears are being stored, and comes face to face with the matriarch parasite. It towers above her at 4 meters tall, and it's grotesquely swollen with hundreds of eggs. Kira senses what to do: she opens all the arks containing the Orbs.
Benny takes Eli to a tunnel of light and tells him to go; Vaughn walks into the light.
The matriarch is taken to a white void where a brown face confronts her; then she appears on a Dominion ship just about to attack DS9; the Jem'Hadar attack.
Akaar demands to know where Kira is, and Ro gives him the runaround. Just as he's about to explode, their badges chirp, and they get word that all the hostage-takers have collapsed; the crisis is over. Akaar says he expects a report from Kira, then beams back to the Trager.
Kassidy pushes and pushes, suffering immeasurable pain, and finally gives birth to a baby girl. Kira shows up just in time--and so does Ben.
A few weeks later: Bajor is due to job the UFP in 26 hours. Kira tells Quark, whose business prospects have fallen through, that his bar has been named the Ferengi Embassy to Bajor, and he the ambassador. He is dumbstruck, and realizes Rom must have saved his bar.
Shar's zhavey asks him to come back to Bajor with her; he tells her he wants to stay and help DS9, and she angrily storms out. He finds he's not all that bothered by it anymore.
Opaka asks to meet with Vedek Yevir. She tells him that Kira is the truest child of the Prophets that's she's ever met, and that it wasn't her, but the Attainder he put on her that caused the spiritual upheaval of Bajor.
Picard sent Ro a security gold uniform as encouragement, but she is still unsure about her future, so she goes to Ziyal's exhibit to think it over. She runs into Garak there, who says he intended to stop by DS9 to see some friends, but that he should be getting back to Cardassia to help his people. Ro decides she's staying, and worries was Quark will think.
Vaughn waits for Prynn by her quarters. When she gets there, he tells her he's never going to stop calling her, and that he doesn't want her forgiveness--he just wants to get to know her. She tears up and says she'll think about it.
Benjamin meets Captain Nerys in her office. She welcomes him back to his office, but he says the office belongs to the commander if the station; Starfleet wants to make him an admiral--but he turned it down. He wants to take his family to the induction ceremony and see where things go from there.
Kira, with her earring back in place, watches the ceremony while holding Odo's hand. She knows he's not staying, but feels certain that they will have a future together. She sees Ambassador Quark standing with Ro in her new uniform. Hiziki Gard is there, having been pardoned, as are Ambassador Word, Chancellor Martok, and Captain Picard. The O'Brien's will be heading to Cardassia after the ceremony where Keiko starts her new job. After the documents are signed, Ben looks at Kira with a "we did it" nod. He is next to his family and new baby girl Rebecca Jae.
Epilogue: Ben enjoys the simple pleasures of home life, knowing that he will be needed soon.
Review: 4 stars. I enjoyed the story, I was glad to see the conspiracy parasites finally get a decent payoff, I liked the Benny/Eli section, etc. The ending did feel a bit rushed though; the parasite queen is easily dispatched, and after reading 7-8 books to see how Sisko would return, well, he just sort of did and that was that. Then the book ended and he didn't even really discuss anything with the other characters.